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1Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Beijing 10081, P. R. China, 2Department of Plant Pathology, China Agricultural University, Beijing 100094, P. R. China1Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Beijing 10081, P. R. China, 2Department of Plant Pathology, China Agricultural University, Beijing 100094, P. R. China